In the "Main mouse wheel" section, if you change (debug-events no) into (debug-events yes), and then start Gimp in a terminal, you should have an inking of what is going on:
Above is for my settings of course, your output would be different.
Code:
Received 'Mouse Wheel Events' (class 'Mouse Wheel')
controller event 'scroll-down-control (Scroll Down (Ctrl))'
(trigger event)
doesn't map to action
Received 'Mouse Wheel Events' (class 'Mouse Wheel')
controller event 'scroll-up-shift-control (Scroll Up (Shift+Ctrl))'
(trigger event)
maps to action 'tools-paint-brush-size-increase-skip'
action was found
Received 'Mouse Wheel Events' (class 'Mouse Wheel')
controller event 'scroll-down-shift-control (Scroll Down (Shift+Ctrl))'
(trigger event)
maps to action 'tools-paint-brush-size-decrease-skip'
action was found
Received 'Mouse Wheel Events' (class 'Mouse Wheel')
controller event 'scroll-down-shift-control (Scroll Down (Shift+Ctrl))'
(trigger event)
maps to action 'tools-paint-brush-size-decrease-skip'
action was found
Received 'Mouse Wheel Events' (class 'Mouse Wheel')
controller event 'scroll-down-shift-control (Scroll Down (Shift+Ctrl))'
(trigger event)
maps to action 'tools-paint-brush-size-decrease-skip'
action was found